Hotkeys: 1-6 for tower placement, S to sell and U to upgrade selected towers, Esc to cancel.
Make combos by placing towers next to each other. All combos have a ?Main? tower (the first one listed in the combo). All the other towers in the combo have to be touching the main tower and all towers must be fully upgraded.
The outer most wall of the tower will not count towards linking. For example: if you drew a square around the tower, to make sure another tower links, it needs to be touching the tower inside of that square.